Standard epidemiological methods to understand and improve Apis mellifera health.

Dennis vanEngelsdorp1*, Eugene Lengerich2, Angela Spleen2, Benjamin Dainat3, James Cresswell4, Kathy Baylis5,  Bach Kim Nguyen6, Victoria Soroker7, Robyn Underwood1, Hannelie Human8, Yves Le Conte9 and Claude Saegerman10.

1 Department of Entomology, 3136 Plant Sciences, University of Maryland, College Park, MD 20742, USA.
2 Department of Public Health Sciences, College of Medicine, The Pennsylvania State University, Hershey, PA, 17033, USA.
3 Swiss Bee Research Centre, Agroscope Liebefeld-Posieux ALP-Haras, Schwarzenburgstrasse 161, 3003 Bern, Switzerland.
4 Biosciences, College of Life & Environmental Sciences, University of Exeter, Hatherly Laboratories, Prince of Wales Road,
Exeter, EX4 4PS, UK.
5 Department of Agriculture and Consumer Economics, University of Illinois, Urbana, IL 61801, USA.
6 Department of Functional and Evolutionary Entomology, University of Liege, Gembloux Agro-Bio Tech, B-5030 Gembloux,
7 Department of Entomology, Agricultural Research Organization, The Volcani Center P.O.B. 6, Bet Dagan 50250, Israel.
8 Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa.
9 INRA, UR 406 Abeilles et Environnement, Site Agroparc, Domaine St Paul, 84914 Avignon Cedex 9, France.
10 Research Unit in Epidemiology and Risk Analysis Applied to Veterinary Sciences (UREAR-ULg), Department of Infectious and
Parasitic Diseases, Faculty of Veterinary Medicine, University of Liège, Boulevard de Colonster 20, B42, 4000 Liège, Belgium.
Received 1 June 2012, accepted subject to revision 26 October 2012, accepted for publication 16 July 2013.

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